Vertical strain partitioning in hot Variscan crust: Syn-convergence escape of the Pyrenees in the Iberian-Armorican syntax

Bryan Cochelin, Dominique Chardon, Yoann Denèle, Charles Gumiaux and Benjamin Le Bayon 1 Géosciences Environnement Toulouse, Université de Toulouse, CNRS, IRD, UPS, CNES, 31400 Toulouse, France 2 BRGM DGR/GSO, BP 36009, 45060 Orléans, France 3 IRD, 01 BP 182, Ouagadougou 01, Burkina Faso 4 Département des Sciences de la Terre, Université Ouaga I Professeur Joseph Ki-Zerbo, BP 7021, Ouagadougou, Burkina Faso 5 Université Orléans, CNRS, BRGM, ISTO, UMR 7327, 1A Rue de la Ferollerie, 45071 Orléans, France

Mots clés : Varisque / Pyrénées / syntaxe / orocline / dôme gneissique / transpression / orogène chaud 1 Introduction In convergent or divergent zones, deformation is unevenly distributed within a given structural level and between contiguous lithospheric layers of contrasted rheology (Grocott et al., 2004).Under brittle conditions, strain is partitioned between strike-slip faults and normal or reverse faults for transtension and transpression, respectively (Tikoff and Teyssier, 1994;Allen et al., 1998;Dokka et al., 1998).Under plastic or ductile regime, strain may be partitioned among shear zones with variable orientations and slip, on the one hand, and lower strain domains of homogeneous strain, on another hand (Gapais et al., 1987;Holdsworth and Strachan, 1991;Holdsworth et al., 2002).The upper and lower crusts may be mechanically coupled to various degrees, in such a way that deformation is differently expressed in each layer while partial structural continuity and kinematic compatibility are maintained between them.The transition between these two crustal layers defines an attachment zone (Tikoff et al., 2002(Tikoff et al., , 2004)).In case of complete decoupling, the attachment becomes a décollement or a detachment.In mature orogens, mechanical coupling between crustal layers is influenced by the fact that the lower crust is hot, partially molten and therefore weak, buoyant and prompt to flow under its own weight (Vanderhaeghe and Teyssier, 2001;Chardon et al., 2009).Under convergence, lateral flow of the weak lower crust interferes with thickening of the upper crust across the attachment, which also acts as a filter absorbing downward thickening of the upper crust, upward flow of the lower crust in gneiss domes, and ascent of lower crustal melts (Chardon et al., 2009(Chardon et al., , 2011;;Bajolet et al., 2015).Under synconvergence lateral flow regimes, lower crust exhumation in gneiss domes is not yet well understood.Some models of gneiss domes formation suggest that decoupling is necessary to form such gneiss domes (the channel flow hypothesis e.g., Rey et al., 2010) but are limited to two dimensions.Other models stress that transcurrent faulting is instrumental to enhance lower crust exhumation (e.g., Le Pourhiet et al., 2012;Roger et al., 2015;Rey et al., 2017) but are imposed on releasing fault steps under purely strike-slip conditions and do not integrate convergence-driven crustal shortening.
The present study explores strain partitioning between the upper crust and the lower crust and its interplay with exhumation of the lower crust during the convergence undergone by the Variscan orogen of SW Europe during latest Carboniferous and Permian times .The chosen case study is the Paleozoic crust exposed in the Pyrenees, which recorded HT-LP regional metamorphism and exposes the relationships between portions of the hot mid-crust and the deep upper crust (Zwart, 1986).The Pyrenees belong to the southern foreland basin of the Variscan orogen that accumulated Carboniferous flysch sediments (Engel, 1984) and that is enclosed in a syntax called the Iberian-Armorican Arc (Matte, 1986;Fig. 1).HT-LP metamorphism of the Pyrenees in a foreland context far from the southernmost HP metamorphic relicts of the Montagne Noire (Fig. 1) appears as an anomaly in the orogen.The proximity of the Pyrenees with the Cantabrian orocline (Fig. 1) that was bent at the core of the syntax between 305 and 295 Ma ( Van der Voo, 2004;Weil et al., 2010;Pastor-Galán et al., 2011, 2016;Gutiérrez-Alonso et al., 2012) further suggests a link between metamorphism, plutonism and deformation in the Pyrenees (ca.320-290 Ma; see below) and orocline closure (Denèle et al., 2014).The second aim of our work is therefore to replace the structural pattern of the Pyrenean crust within the evolving kinematic frame of the Variscan orogen, with an emphasis on the closure of the Cantabrian orocline in an abnormally hot foreland setting.The present study is based on an extensive compilation of fabric data and new structural observations at the scale of the Pyrenees that allowed building an integrated regional structural map.A 3D crustal coupling model is proposed to explain the interplay of lower crustal flow and upper crustal transpression as well as oblique (transtensional) extension allowing for the exhumation of gneiss domes under convergence.We show how such a 3D orogenic flow mode recorded the closure of the Cantabrian orocline in a context of hot and thin lithosphere.

Variscan Pyrenees: geological outline and earlier works
The present day Pyrenees form a N100°E trending mountain belt (Fig. 2) that results from the interactions and ultimate collision between the Iberian and Eurasian plates, which took place from the late Cretaceous to the Paleogene (Choukroune, 1992).During convergence, Mesozoic series and their Paleozoic substrate have been shortened to form an asymmetric doubly verging orogenic prism (Choukroune et al., 1989).Paleozoic basement outcrops are found in two main structural domains separated by the North Pyrenean Fault (Fig. 2).The Axial Zone (south of the fault) forms the uplifted core of the orogen and represents the largest continuous basement outcrop, whereas the Basque and North Pyrenean massifs are smaller basement units emerging from their Mesozoic cover North of the fault (Fig. 2).The basement preserves ubiquitous Variscan structures, penetrative strain and metamorphic patterns that have only locally been modified by Meso-Cenozoic faulting and block tilting (Carreras and Debat, 1996;Carreras and Druguet, 2014).The Paleozoic crust of the Pyrenees is characterized by a strong structural contrast between an upper crustal level, called the superstructure, and the roof of a lower-crustal level outcropping in metamorphic domes, called the infrastructure (de Sitter and Zwart, 1960;Zwart, 1979;Carreras and Capella, 1994).The superstructure is made of low-grade Ordovician to Carboniferous metasedimentary rocks.They are intruded by large plutons of calc-alkaline affinity comprising gabbros, tonalites, granodiorites and granites of mantle or hybrid sources (Guitard et al., 1984;Debon et al., 1996;Roberts et al., 2000).The infrastructure has undergone HT-LP metamorphism (Guitard et al., 1996).It comprises Ediacarian-early Cambrian paragneisses hosting Ordovician granites (470-450 Ma) turned into orthogneiss sheets, as well as Ordovician micaschists and paragneisses (Carreras and Druguet, 2014;Denèle et al., 2014, and references therein).Magmatic rocks of the infrastructure are crustally derived peraluminous granites (mostly sills) and associated migmatites, as well as kilometerscale sills of diorites and gabbros.The transition between the shallow fabrics of the infrastructure and the steep fabrics of the superstructure roughly coincides with the Andalusite isograd that underlines the map shape of the domes (Guitard et al., 1996;Debon and Guitard, 1996, e.g., Fig. 2).Attenuation of the isograds at the roof of the infrastructure results in a strong field metamorphic gradient (Zwart, 1979;Wickham andOxburgh, 1985, 1986;Van den Eeckhout and Zwart, 1988;Paquet and Mansy, 1991;de Saint-Blanquat, 1993;Guitard et al., 1996;Vilà et al., 2007;Mezger and Régnier, 2016).Clockwise PTt paths in the infrastructure have been documented from peak pressures of ca.4.5 kb with significant heating during decompression and later cooling along a steep geotherm (Mezger and Passchier, 2003;Mezger et al., 2004;Mezger, 2005;Vilà et al., 2007).Recently, Aguilar et al. (2015) reported metamorphic relicts suggestive of an early hightemperature peak pressure of 7.5 Kbar in the Roc the France dome (Fig. 2).Pre-main cleavage, folds and thrust faults have been reported from the superstructure (Mey, 1967(Mey, , 1968;;Valero, 1974;Müller and Roger, 1977;Zwart, 1986;Speksnijder, 1987;Carreras and Capella, 1994;Matte, 2002).Pre-main cleavage recumbent fold nappe models of the infrastructure have been proposed by Vilà et al. (2007) and Aguilar et al. (2015) for the Roc de France and Albères massifs (Fig. 2).Such models recall early interpretations of fold interference patterns emphasizing early recumbent folding (i.e., Penninic style of folding) and later superimposition of upright structures (Guitard, 1964;Séguret and Proust, 1968;Guitard et al., 1984Guitard et al., , 1998;;Carreras and Capella, 1994).
Calc-akaline plutons of the superstructure yielded U-Pb zircon ages of 310 to 290 Ma (Aguilar et al., 2014;Denèle et al., 2014 and references therein;Druguet et al., 2014).Mezger (2010) and Mezger and Gerdes (2016) obtained ∼340 Ma ages for the peraluminous granites of the Aston dome (Fig. 2) using LA-ICP-MS on zircons, at odd with ages of ∼306 Ma obtained on the comparable nearby intrusions with the same method by Denèle et al. (2014).Those later authors interpreted the ∼340 Ma (Visean) ages as that of an inherited crustal component in those granites instead of that of their magmatic crystallization.Esteban et al. (2015) distinguished two U-Pb zircon age groups in the Lys-Caillaouas pluton (Fig. 2) intruding the roof of the infrastructure at 307 ± 3 Ma and 300 ± 2 Ma, and suggested that the first group dated metamorphism and the second pluton crystallization.Recently, Kilzi et al. (2016) obtained an age of 294 ± 1 Ma by LA-ICPMS on zircons in a mantle-derived dioritic sill hosted by migmatites of the Gavarnie massif (Fig. 2).The oldest recognized map-scale folds (Matte, 2002) affect lower to middle Pennsylvanian (318-307 Ma) flysch deposits (Delvolvé et al., 1993), providing a maximum age constraint on deformation of the superstructure.This, together with dating of synkinematic HT metamorphism, melting, plutonism and cooling from high temperature in the infrastructure and the ages of synkinematic plutons in the superstructure (310-290 Ma), suggests that Variscan deformation in the Pyrenees took place between ca.320 and 290 Ma and between ca.310 and 290 Ma mostly under HT-LP conditions.
Most authors agree on the fact that the steep structures and fabrics of the superstructure have recorded N-S shortening.However, the tectonic significance of the deformation patterns in the infrastructure and their spatial and temporal relationships with those of the superstructure have been debated (Carreras and Capella, 1994).Coeval deformation of the superstructure and the infrastructure under N-S shortening regime was favored by Zwart (1979Zwart ( , 1986)), as well as by Soula (1982), Soula et al. (1986a, b) and Pouget (1991), who suggested diapiric emplacement of the domes.Attenuation of the isograds and associated high thermal gradient point to an extensional component of deformation at the Infrastructure -Superstructure transition (Wickham andOxburgh, 1985, 1986;Van den Eeckhout and Zwart, 1988;de Saint-Blanquat, 1993).In the Axial Zone, extension was interpreted to have taken place before (Soula et al., 1986a), during (Mezger and Passchier, 2003) or after (Van den Eeckhout and Zwart, 1988;Vissers, 1992) shortening of the superstructure.In the eastern Pyrenees, HT deformation and magmatism have been shown to be coeval with regional-scale dextral-reverse shear zones (Carreras and Capella, 1994), in agreement with synkinematic emplacement of the plutons in the superstructure in an dextral transpressional context (Bouchez and Gleizes, 1995;Gleizes et al., 1998aGleizes et al., , 2001Gleizes et al., , 2006;;Aurejac et al., 2004;Carreras et al., 2004;Román-Berdiel et al., 2004, 2006).Recent structural analysis of the Aston, Hospitalet and Bossost domes (Fig. 2) led to models of lateral flow of the infrastructure and dome formation during regional transpression (Mezger and Passchier, 2003;Denèle et al., 2007Denèle et al., , 2009;;Mezger, 2009).

Methodology and systematics of the structural markers
We have compiled the planar and linear strain fabrics, structures (shear zones and faults) and associated kinematic criteria affecting the Paleozoic rocks of the Pyrenees.Structural data come from structural studies of the literature as well as from the French and Spanish geological maps.Evaluation and validation of the data as well as gap filling in the sampling were made through field surveys over key areas at 1873 stations, most of the kinematic measurements having been acquired for the present study.Harmonization of the data over a total of 16 551 stations led to a map of the Variscan structural and kinematic patterns at the scale of the Pyrenees (Fig. 3), which is complemented by a series of regional cross-sections through the Axial Zone (Fig. 4).Given their areal extent, the Axial Zone and  and Guitard (1996) for the corresponding regional-scale isograd map pattern.the Basque Massifs are the focus of our work.Information on the North Pyrenean Massifs has been incorporated to the structural map mostly from the literature (Fig. 3).Analysis of the regional strain field is complemented by studies at the scale of two metamorphic domes (Bossost and Lys-Caillaouas; Fig. 2).These studies allow for the documentation of the deformation of the infrastructure, its interference patterns with the superstructure, and dome emplacement processes in greater details.
The main and ubiquitous penetrative planar fabric affecting the superstructure (called hereafter Ss) is referred to as a schistosity, which is best expressed in the dominant argilous and pelitic lithologies of the low-grade metasediments.Depending on the strain intensity and the lithology, that fabric may become a foliation in recrystallized carbonates, more commonly a fracture cleavage (in competent sandstones or carbonates) or a slaty cleavage (e.g., in calcschists).Ss is steeply dipping and is generally axial planar to macroscopic to regional-scale, steep to upright folds that range from open to isoclinal.Ss schistosity commonly bears a steep stretching and/ or mineral lineation (Ls), which is marked mostly by elongated quartz aggregates, calcite ribbons, or white mica flakes.The main penetrative planar fabric of the infrastructure is a foliation (called Si) bearing a mineral-stretching lineation (Li) defined by quartz ribbons, elongated feldspars or feldspar aggregates, fibrolitic sillimanite or aligned mica flakes or aggregates.Regional-scale schistosity (Ss) and foliation (Si) trajectories are drawn on Figure 3 together with lineation (Ls and Li) measurements, as well as regional shear zones revealed by fabric trajectories and field mapping.Foliation trajectories in the plutons were drawn on the basis of the published data.Kinematic information related to non-coaxial macroscopic deformation features such as C' shear zones and shear bands, asymmetrical boudinage and sigma clasts are also reported on Figure 3 for the infrastructure and the superstructure and the regional shear zones (see below).At a number of stations in the infrastructure, symmetrical boudinage or a comparable number of contradictory shear criteria of the same type are observed for a common stretching direction.Such a configuration is interpreted to reflect local bulk coaxial deformation (Fig. 3a).
Figure 5 synthetizes the fabric data measurements.Field illustrations of fabric and kinematic patterns are illustrated on Figures 6 and 7 for the superstructure and the infrastructure, respectively.A pre-Ss cleavage is preserved in lower strain domains of the superstructure (Figs. 2 and 3).This cleavage is associated with the early map-scale folds and has been mapped   as S1 (Fig. 3a).In the present contribution, post-Paleozoic faults, fabrics and structures are referred to as "Alpine".

Structural and kinematic analysis 4.1 Large-scale patterns
Ss schistosity trajectories of the superstructure trend N90-N100°E and define, together with an anastomozed network of regional shear zones, the structural grain of the Axial Zone (Fig. 5a).The shear zone network mostly consists in two sets.Shear zones of the first set trend dominantly E-W, whereas those of the second set preferentially trend N130°E.A large part of the shear zone network had been mapped by Carreras (2001) in the eastern Axial Zone and our map shows that it extents further to the West (Fig. 3).Low-strain lens-shape domains in between the shear zones are up to 50-km long and 10-to 20-km wide and commonly host the plutons and the metamorphic domes (Fig. 3).The shear zones are steep, kilometer-thick, retrograde, amphibolite to greenschist facies mylonite zones (e.g., Carreras, 2001;Fig. 4).With the exception of the Esera Gistain shear zone South of the Lys-Caillaouas dome (Figs. 2 and 3; see below), the regional shear zones are systematically characterized by moderately to steeply West-plunging stretching lineations and reverse-dextral kinematics (Fig. 6d and e).The superstructure in the low-strain domains displays homogeneous deformation marked by steep rectilinear schistosity trajectories (Fig. 6a) and steeply plunging stretching lineations (Fig. 5b).Steep stretching is further attested by decameter to meter-scale C' shears bearing down-dip striations with top-to-the south sense of shear throughout the southern half of the Axial Zone (Figs. 3, 5b and 6b).In the northern half of the Axial Zone, both North-side-up (mostly) and South-side-up C' shears affect the steep schistosity (Figs. 3  and 6c).In the vicinity of plutons and especially in their contact aureoles, Ls stretching lineations acquire a shallower plunge and C' shear bands and shear zones have dextral senses of movement (Figs. 3 and 5b).The plutons exhibit dominantly concentric foliation trajectories and schistosity triple points in their country rocks near their extremities (Marre, 1973;Aparicio, 1975;Messaoudi, 1990;1998b;Debon et al., 1996).Some of the plutons are bounded by the reverse-dextral mylonitic shear zones described above (Figs.3, 6d and e).Those pluton-bounding shear zones typically host C/S fabrics, i.e., non-coaxial shearing features of syn-kinematic cooling granite as defined by Gapais (1989) (Têt, Perthus and Mérens shear zones; Figs.2-4).In the southern part of the Axial Zone, from steep, Ss becomes shallowly dipping going southward (Fig. 4, sections 2-5) and the Ls stretching lineations become shallower accordingly (Fig. 5a and b).The resulting upward fan of Ss schistosity underlies the basal unconformities of the Permian or Meso-Cenozoic strata, which dip moderately (40°) to the south (Fig. 4, cross-section 3).As earlier argued by Carreras and Debat (1996), post-Paleozoic southward tilting and bending of those unconformities explain the flexuring of their substrate that produce the Ss fan.Restoration of the unconformities back to the horizontal shows that Ss was originally North-dipping around at least 60°before Meso-Cenozoic deformation (Carreras and Debat, 1996).Therefore, the schistosity was consistently steep to vertical throughout the superstructure by the end of the Paleozoic.
Metamorphic domes are windows onto the infrastructure.They have an elliptical map shape, with their great axis parallel to the regional structural grain (Fig. 3).They are of two types.Eastern domes (e.g., Canigou, Aston, Hospitalet) mostly expose Ordovician orthogneiss sheets that usually form their envelope, whilst western domes (e.g., Bossost, Lys-Caillaouas, Chiroulet, Lesponne) are devoid of orthogneisses (Figs.2-4).The roof of the infrastructure corresponds to the upper contact of the orthogneisses with schistose rocks of the superstructure for the eastern domes.That roof lies within and preferentially near the top of Ordovician series for the western domes (Fig. 4).The eastern domes exhibit steep and locally southerly overturned limbs that are attenuated by the regional reversedextral shear zones (Denèle et al., 2007;Vilà et al., 2007; Figs. 2 and 4).Western domes such as the Bossost dome are less amplified and not clearly asymmetrical (Fig. 4, cross-section 2).Si foliations are dominantly shallowly dipping and the scattering of their poles on a N-striking vertical plane (Fig. 5c) attests to the E-W trend of the domes hinges (Fig. 3).
An overwhelming part of the Li lineations (90%) in the Axial Zone trend N70 to N120°E, i.e., at a very low angle to the schistosity trajectories of the superstructure (Figs.3b and 5d).Hereafter, they are therefore called longitudinal Li lineations.They are shallowly plunging and acquire moderate to steep plunges on domes' flanks.N90 to N120°E trending Li lineations are dominant over most of the Axial Zone, whereas N70 to N90°E trending Li lineations are confined to the Canigou and Roc de France Massifs (Figs. 2 and 3).At the scale of some metamorphic domes (e.g., Canigou, and southern Bossost dome), Li trajectories have sigmoidal map trajectories showing a transition from longitudinal (domeparallel) flow at the core of domes to oblique stretching on domes' envelops (Fig. 3).
Si fabrics are affected by discreet to pervasive C' shear zones and shear bands separating asymmetrical boudins and sigma-type porphyroclasts.Those shears bear lineations trending parallel to Li in the host rock and are commonly filled with fabric-free granitic material collected from the host migmatitic gneisses, attesting to the HT, melt-present conditions of non-coaxial longitudinal shearing of the infrastructure (e.g., Fig. 7a).A large number of Li stations (50%) reveal top-to-the E or SE sense of shear, whereas top-tothe W or NW kinematics are documented at 31% of the Li localities (Figs. 3 and 7b).Stations with coexisting shear criteria of opposite sense and/or symmetrical boudins (Fig. 7c) represent 19% of the data (Fig. 3).The Aston and Hospitalet domes display preferential eastward shearing and others like the Chiroulet and Canigou are dominated by westward shearing (Figs. 2,3,7a and b).Longitudinal shearing also occurs in shear zones flanking the lateral termination of some domes (e.g., Hospitalet, Canigou, Figs. 2 and 3).At these locations, asymmetrical shear bands and shear zones evolve from melt-present, amphibolite-facies fabrics to greenschist-facies fabrics and strain localization, indicating syn-cooling shearing.
Li mineral-stretching lineations trending at a high angle to the structural grain (called hereafter transverse Li lineations) represent 10% of the measurements and are restrained to domes' envelop (Canigou, Aston and Chiroulet massifs; Figs. 2  and 3).Transverse lineations are associated with hundreds of meter to kilometer-thick retrograde, syn-melt (Fig. 7d) to greenschist-facies shear zones flanking the domes.Those shear zones have systematic normal-sense of shear.
In most domes, there is a gradient from longitudinal to transverse lineations, going from domes' cores to domes flanks, whereas in parts of specific domes (e.g., Aston, Canigou, Figs. 2 and 3) longitudinal and transverse Li lineations coexist.Given that both lineations are marked by HT minerals or mineral aggregates of leucosomes, they must have overlapped in time under HT and partial melting conditions.Be they associated with longitudinal, oblique or transversal stretching, the dome-flanking shear zones display systematic normal component of shearing and are retrograde, with shearing features going from melt-present or amphibolite facies metamorphic assemblages to brittle-ductile.This, together with the pattern of attenuated isograds around domes (e.g., Gibson and Bickle, 1994;Mezger and Régnier, 2016) indicates that extension contributed to dome exhumation.The study of the Bossost and Lys-Caillaouas domes (see Sect. 4.2 and 4.3 below) allows illustrating in greater details extensional dome emplacement modes as well as the strain interference patterns between domes' fabrics and those of the superstructure.
Metamorphic, strain and kinematic patterns preserved north of the North Pyrenean Fault (i.e., North Pyrenean and Ursuya massifs; Figs. 2 and 3) are distinct from those of the Axial Zone.Their fabrics may not be readily compared to those of the Axial Zone given the rotation the Axial Zone must have undergone in the Mesozoic with respect to areas located today north of the North Pyrenean Fault.However, for a purpose of convenience in the description of the structural pattern, the Axial Zone terminology is used here to refer to North Pyrenean fabrics.All the North Pyrenean massifs preserve granulitefacies rocks in their infrastructure.N-S stretching and shearing is pervasive in the Agly, St Barthélémy and Castillon massifs (Figs. 2 and 3).Shearing in the Agly and St Barthélémy massifs has been shown to result from extensional exhumation of the infrastructure and strain localization in detachments at its top (Bouhallier et al., 1991;de Saint-Blanquat, 1993).Interestingly, the Ursuya massif bears longitudinal Li in its infrastructure, whereas extensional shearing and detachment faulting was activated by top-to-the South (i.e., transverse) kinematics (our own observations; Figs. 2 and 3).This shows a kinematic contrast between deep stretching and higher-level shearing, comparable to the Canigou and Aston domes (Figs. 2  and 3).

The Bossost dome
In contrast with other domes of the Axial Zone, the Bossost dome shows relationships with the surrounding rocks of the superstructure that have not been obscured by the regional reverse-dextral shear zones.The dome has only been affected by localized post-Paleozoic faulting that has cut the dome in two sub-domes (Figs. 3, 4 and 8).The andalusite isograd roughly coincides with the outer concentric trajectories of the shallowly to moderately dipping foliations of the infrastructure (Si) and the steeper schistosity of the superstructure (Ss), with the noticeable exception of the southern flank of the southern subdome where the isograd rises up into the steep fabrics and folds of the Devonian sedimentary pile (Fig. 8a).The northern sub-dome exposes a deep portion of the northern flank of the original dome, whereas the southern sub-dome exposes mostly the E-W elongate core and the southern flank of the original dome.Peraluminous granite sills are intercalated in the dome, particularly in its northern flank, which display mineralstretching lineations (Li) of two main trends (see also Mezger and Passchier, 2003).The dominant lineation is longitudinal, trending parallel or at a low angle to the map axis of the dome and a few transverse lineations are documented in the vicinity of the northern margin of the dome, sometime on the same outcrops than the longitudinal Li (see also Mezger and Passchier, 2003).The fact that both longitudinal and transverse lineations are underlined by the same HT minerals suggests flattening strain in this part of the dome.The southern subdome exhibits overall sigmoidal Li trajectories going from longitudinal and N100°E trending in its core to transverse (up to N160À170°E trending) on its flanks (Fig. 8b).Top-to-the NW kinematics is documented from the northern limb and pervasive top-to-the SE or SSE shearing affects the southern flank.In both cases, shear criteria are towards the azimuth of plunge of Li lineations.
Top-to-the SE shearing is best expressed in a ca. 2 km-thick shear zone (Fig. 9) exposed between Arres and Benos, which affects both the paragneisses and the overlying Devonian schists and marbles (Figs.8a and 9; Mezger and Passchier, 2003).Shearing features are numerous, going from asymmetrical porphyroclast systems to extensional C' shear zones and shear bands having contributed to thinning of the shear zone (Fig. 7e).Furthermore, shear zones and shear bands go from sillimanite-stable to brittle-ductile, indicating cooling of the rock pile during non-coaxial shearing and vertical thinning compatible with extension along the dome's flank.The extensional shear zone is crosscut by a steep mylonite zone localized in the Silurian black shales (Fig. 9).That mylonite zone fans up upward and northward to mold the domal envelop of the paragneisses and form an array of meter-to decameterscale, asymmetrical and anastomosed shears (Fig. 10a) with south-side up or top-to-the north kinematics, whereas northside up shears are also documented near the Ordovician-Silurian contact (see also García-Sansegundo, 1990, 1992, Fig. 10b).A detailed cross-section of this sector of the dome's flank is provided in Figure 10.Above, Devonian series are affected mostly by steep south-dipping thrusts rooted in the Silurian shales as well as steep folds with an axial planar schistosity corresponding to Ss, i.e., the regional planar fabric of the superstructure (Fig. 9).Ls lineation in the superstructure is systematically down-dip on Ss.Near the Andalusite isograd, micaschists with shallow Si foliation are affected by micro to meter-scale, E-W trending upright folds with an axial planar cleavage.Over a few hundreds of meters going up in the structural pile, such folds become tighter up to transposition of Si by the steep Ss schistosity.Along this strain gradient, folded boudins are common, further attesting to the reworking of the dome-flanking shear zone by the steep fabrics of the superstructure (Ss and Ls).
The structural and kinematic configuration of the southern flank of the dome as reported above is symmetrical with respect to that of its northern limb.There, the shallowly Ndipping Si foliations of the dome are blanketed by the Silurian shales onto which north-dipping thrust faults branch.These faults affect, and are kinematically coherent with, the E-W upright folds and associated Ss schistosity of the overlying Devonian series (Fig. 9).The crestal region of the dome displays fabric interference patterns formed by vertical shortening and longitudinal stretching (Si/Li), on one hand, and N-S shortening and vertical stretching (Ss/Ls), on another hand (Fig. 9).Such interference patterns are interpreted to reflect a longitudinal schistosity triple point along the dome's crest.
Exhumation of the infrastructure into the Bossost dome evolved from deep dominant longitudinal flow (as preserved in dome's core) to oblique, dominantly SE-directed, syn-cooling stretching and strain localization along extensional shear zones flanking the dome.Low-grade rocks of the superstructure preserved above the extensional contact of the dome exhibit penetrative fabric and fold patterns produced by N-S shortening, vertical stretching and reverse faulting on dome's flank towards the dome's crest.Shortening and thrusting recorded by the superstructure outlasted extensional exhumation of the dome, whereas no penetrative steep fabrics are preserved in the exposed infrastructure.This, together with the interference patterns developed between the dome's roof and the superstructure, suggests that dome amplification and regional shortening have overlapped in time.Field relations between the infrastructure and the superstructure as reported here for the Bossost dome are comparable to those we have examined around other domes in areas where the relationships with the superstructure have not been modified by the regional reverse-dextral shear zones (Chiroulet, Albères and Canigou domes; Figs. 2 and 3, see also Fig. 7f).Those observations argue for a common structural and kinematic framework of lower crustal flow, extensional exhumation and shortening patterns at the scale of the Axial Zone.

The Lys-Caillaouas dome
The Lys-Caillaouas massif exposes the southern limb of a N130°E-trending dome (Figs.2-4, cross-section 2).The dome is made of paragneisses (locally migmatitic) hosting the Lys-Caillaouas pluton, which consists of a dioritic core surrounded by granites (Clin et al., 1986;Hilario-Or us, 2004) that ultimately crystallized at 300 ± 2 Ma (zircon SIMS U-Pb date of Esteban et al., 2015).The pluton is a sill interleaved within the paragneisses that roots into the Esera-Gistain shear zone, which truncates and bounds the dome to the South (Fig. 11a).Shallowly plunging mineral-stretching lineations (Li) in both the paragneisses and the plutonic rocks are dominantly WNW trending in the infrastructure (Fig. 3).
Bulk coaxial longitudinal deformation attested by opposite shear criteria and symmetrical boudinage affects the pluton's country rocks (Fig. 7c), whereas ESE-directed shearing characterizes the rest of the infrastructure (Figs. 3  and 11a).A southward strain gradient into the southern boundary shear zone is illustrated by fabrics evolving from sub-magmatic to mylonitic, indicating syn-cooling strain localization.Initially isotropic diorite is affected by typical C/ S fabrics (Gapais, 1989), whereas micaschists and paragneisses of the host rocks and the pluton's septa display systematic meter-scale C' shear zones and shear bands of sinistral-normal kinematics (top-to-the ESE) on progressively steeper foliations going southward (Fig. 11b and c).Furthermore, the southern boundary shear zone marks a metamorphic break between the Sillimanite paragneisses in its footwall and very low-grade sediments in its hangingwall.The asymmetrical map shape and truncation pattern of the pluton by the shear zone and the asymmetrical fabric trajectories against the shear zone further argue for apparent sinistral sense of movement of the shear zone.The Esera-Gistain shear zone is therefore interpreted as a left-lateral transtensional shear zone that contributed to exhumation of the dome under .Given the 307 ± 3 Ma SIMS U-Pb age of metamorphic zircons of the plutonic complex (Esteban et al., 2015) and the final crystallization age of the pluton (300 ± 2 Ma), longitudinal stretching and transtensional exhumation must have been active since ca.310 Ma and continued until after 300 Ma, considering that shearing occurred in the transtensional shear zones until after cooling from high temperature.

Early Variscan deformation
Pre-Ss map-scale folds, cleavage (S1) and faults are best observed near the Western termination of the Axial Zone (Mirouse, 1962;Valero, 1974;Müller and Roger, 1977;Matte, 2002) (Fig. 3).In this area, shallower levels of the superstructure are exposed given the differential longitudinal exhumation of the Axial Zone, which allowed a westward-increasing preservation of post-Ordovician stratigraphic pile westward (Fig. 4).Ss/Ls fabrics are less intense than in the rest of the Axial Zone and allow preservation of D1 features, whose kinematic significance is B. Cochelin et al.: BSGF 2017, 188, 39 unambiguously interpreted, as opposed to locations further to the East (Guitard et al., 1984(Guitard et al., , 1998)).Over the area south of the Eaux-Chaudes pluton (Fig. 2), Matte (2002) reported southwestward to westward thrusting on the basis of overturned to recumbent fold asymmetries and F1 folds -S1 cleavage relationships.We report here on a cross-section made at the western tip of the Axial Zone (Figs. 3 and 12).Devonian carbonates and overlying late Carboniferous flysch series are involved in km-scale folds inclined to slightly overturned towards the west (Fig. 12a).Folds short limbs are affected and attenuated by E-dipping thrust faults.In the field, such faults typically show down-dip striations as well as top-to-the West reverse shear senses, consistent with microfolds' asymmetry (Fig. 12b).Such a structural pattern, which was also described by García-Sansegundo (1992, p. 251), is observed in the Ordovician-Devonian series of the Basque Massifs as well despite the occurrence of a younger E-W Ss cleavage (Figs. 2 and 3) (Valle de Lersundi et al., 1972;Villalobos Vilches and del Valle de Lersundi, 1973 and our own observations).Such geometries and kinematics indicate that the structural style of early Variscan (D1) deformation was that of a west-vergent fold-and-thrust belt in the westernmost Axial Zone and the Basque Massifs.Whether D1 fold-and-thrust relicts found elsewhere in the Axial Zone (Fig. 3) were originally N-S and rotated to roughly E-W by the main deformation that produced Ss is not known.D1 folds and thrusts may as well have originally been part of a virgation going from ESE-trending in most of the Axial Zone to N-trending in the westernmost Axial Zone and Basque Massifs.

Late-and Post-Variscan deformation
In the Basque Massifs, a N-striking fracture cleavage is superimposed on Ss over an area coinciding with a regional gravity anomaly surrounding the 267 Ma Aya pluton (Figs. 2 and  3; see Debon and Guitard, 1996).This cleavage being crosscut by the Alpine cleavage affecting the neighboring Triassic deposits (Villalobos Vilches and del Valle de Lersundi, 1973; our own observations), it is most likely Permian in age and linked to the emplacement of the Aya pluton (Denèle et al., 2012).
In the Southern part of the Axial Zone, Alpine thrust faults root into the steep Variscan shear zones and acquire shallower northern dip towards the surface.These faults achieved thrusting of basement over the Meso-Cenozoic series of the Southern Pyrenean zone, with cumulated throw ranging from a few kilometers to 10-15 km (e.g., the Eaux-Chaudes and Orri thrusts; Fig. 4, cross-sections 2 and 3, see also Déramond, 1979;Majesté-Menjoulas, 1979).

Structural synthesis
The consistent orientation of Ss/Ls fabrics of the Axial Zone define a homogeneous strain field reflecting N-S horizontal shortening and dominantly steep stretching (i.e., bulk vertical pure shear) with a subsidiary component of simple shear (as attested by dominant north-side-up kinematics in the southern Axial Zone).The map symmetry of the regional anastomozed shear zone pattern with respect to  10.(a) Detailed cross-section of the transition between the infrastructure and the superstructure interpreted as an attachment (southeastern limb of the Bossost dome, road cut NW of Arres, Fig. 8a).(b) Structural elements measured along the cross-section (lower hemisphere, equal area projection).Schistosity and associated shears of the superstructure (in black) are superimposed on the foliations molding the roof of the infrastructure (in red).The steep reverse shears are both North-and South-verging and branch onto the roof of the infrastructure.By contrast, the regional transpressive shear zones cut across the attachment (Fig. 4).rectilinear trajectories of steep Ss is remarkable, with the schistosity bisecting the angle between the two sets of shear zones (Fig. 3b).Such a symmetry is indicative of a kinematic coherency between, and coeval activation of, the regional schistosity and the shear zones in response to bulk inhomogeneous deformation (Gapais et al., 1987(Gapais et al., , 2005)).This deformation was partitioned between bulk pure sheardominated deformation in low strain lenses and non-coaxial strain localization in the reverse-dextral (i.e., transpressive) shear zones.The regional pattern of bulk inhomogeneous dextral transpression (Fig. 3b) is also consistent with structural patterns documented at various scales throughout the Axial Zone and particularly those produced by synkinematic pluton emplacements (Carreras and Capella, 1994;Gleizes et al., 1997Gleizes et al., , 1998b;;Denèle et al., 2007Denèle et al., , 2009;;Carreras and Druguet, 2014).
Pervasive longitudinal stretching and shearing of the infrastructure attest to lateral flow on a regional scale.Lateral flow of the infrastructure is kinematically compatible with transverse shortening of the superstructure.Furthermore, longitudinal flow of the infrastructure and shortening deformation in the superstructure overlapped in time for at least ca.15 Myr.Indeed, syn-kinematic HT metamorphism, partial melting and sill emplacement in the infrastructure lasted from at least ∼310 to 294 Ma (possibly up to ca. 320 Ma; e.g., Aguilar et al., 2014), whereas bulk inhomogeneous transpression and syn-kinematic pluton emplacements in the superstructure lasted from ∼310 to 290 Ma.This geochronological overlap is also documented at the scale of specific domes/plutons such as the Lys-Caillaouas dome (Esteban et al., 2015), La Jonquera pluton (Aguilar et al., 2015) and the Aston dome (Denèle et al., 2009(Denèle et al., , 2014; Fig. 2).
The emplacement mode of the gneiss domes of the Axial Zone attests to the extensional exhumation of the ductile buoyant infrastructure in the lower superstructure.Although they ultimately evolved under retrograde metamorphic This indicates that a dominant transtensional (i.e., oblique) component of stretching was permitted under convergence to allow exhumation of the infrastructure.This transtensional component of stretching is also indicated by the obliquity of longitudinal Li lineations on the domes' hinges (comparison of Fig. 5c and d).The dominance of southeastward-directed extension requires a favorable boundary condition to the east or southeast of the Pyrenees (in present-day coordinates).
Potential (early) thickening history of the infrastructure has been erased by lateral flow and extension at its roof.Although interference strain patterns may argue for local multistage deformation histories, superimposition of distinct and diachronic regional Variscan deformation phases cannot be substantiated at the scale of the Pyrenees from 310 Ma onward (i.e., after D1 fold-and-thrust deformation; see also Carreras and Capella, 1995 in their reply to Aerden, 1995).Our conclusions concur with those of Mezger and Passchier (2003), who argued in their study of the Bossost dome for local extension permitted under regional shortening regime.Our work further allows addressing the causes and consequences of coeval orogen-parallel lower crustal flow, extensional doming and orogen-normal shortening at the scale of the Variscan Pyrenees.

Discussion
6.1 Three-dimensional flow mode of the Variscan Pyrenean crust Coeval transpression of the superstructure and lateral flow of the infrastructure typically compare to those of ultra-hot orogens, indicative of a particularly weak lithosphere submitted to convergence (Cagnard et al., 2006;Cruden et al., 2006;Chardon et al., 2009).The transition zone between the infrastructure and the superstructure as originally defined by de Sitter and Zwart (1960) represents an attachment (Chardon et al., 2009;Figs. 10 and 13a).The role of the attachment was to maintain vertical strain continuity through the crust by monitoring the conversion of orogen-normal shortening into vertical stretching and lateral flow between the superstructure and the infrastructure, respectively.The attachment further had to allow the rise of the partially molten buoyant infrastructure into transtensional domes (Fig. 13a).The reason why transtension, instead of orogen-parallel extension, enhanced dome emplacement is that it combines eastward lateral escape (orogen-parallel flow end-member) and gravitational collapse (orogen-normal end-member)  (Chardon et al., 2011).The attachment also had to absorb differential horizontal and vertical displacements due to the contrasted deformation modes of the lower and upper crusts that imply spatially and temporally contrasted transversal contraction and lateral elongation.We interpret transtensional dome emplacement and slip on the regional transpressive shear zones to have absorbed kinematic incoherencies due to those contrasts.Although the attachment's thickness and kinematics evolved through space and time, its roof seems to roughly mark an isotherm (i.e., the andalusite isograd), which must have dictated a rheological threshold in the crust.
Transtensional doming into a thickening superstructure led to limited finite exhumation so that infrastructure rocks were not brought to the surface and the domes are not flanked by extensional sedimentary basins.Therefore, they are not metamorphic core complexes as defined in the Basin and Range province (e.g., Davis and Coney, 1979;Lister and Davis, 1989).We suggest that they represent syn-convergence transtensional gneiss domes, which may be common in thermally mature convergent orogens and have been mistaken for metamorphic core complexes or compressive or diapiric domes.
Shortening deformation of the superstructure outlasted fabrics of the exhumed roof of the infrastructure.Likewise, the regional transpressive shear zones reactivate, or are superimposed on, the extensional shear zones flanking the domes (e. g., Canigou dome, Figs. 2 and 4).Furthermore, domes have been amplified by N-S shortening and locally overturned by reverse-dextral slip on the regional shear zones (Vilà et al., 2007;Denèle et al., 2007Denèle et al., , 2009;;Figs. 4 and 13b).The regional transpressive shear zones crosscut the attachment (Figs. 4 and  13b) and were active until after ca.290 Ma, i.e., the age of the youngest pegmatites they affect (Druguet et al., 2014;Van Lichtervelde et al., 2017).At least some of them must have been through-going during earlier (310-300 Ma) crustal flow of the infrastructure because they acted as magma channels at the time (Denèle et al., 2008, Fig. 13b).Furthermore, synkinematic mineral assemblages in the shear zones attest to cooling during strain localization.These considerations argue for increasing coupling across the attachment during crustal cooling and stiffening of the infrastructure after ca.295 Ma (i.e., the latest cooling age from high temperature of the infrastructure).

Implications for early Variscan and post-Variscan deformation of the Pyrenees
Despite intense deformation of the superstructure where upright isoclinal folds are common, the envelope of the stratigraphic markers and the metamorphic isograds remained roughly horizontal (Fig. 4).This precludes the occurrence of Variscan crustal-scale thrusts that would have disrupted the stratigraphic pile and produced major metamorphic breaks.Therefore, D1 deformation must have involved thin-skinned instead of thick-skinned fold-and-thrust tectonics.Likewise, early (pre-main Ss cleavage) recumbent fold nappes involving the infrastructure are also unlikely in the Pyrenees.
Maintenance of a flat envelop for the Paleozoic sedimentary pile and the isograds over the Axial Zone also suggests that the occurrence of Alpine thrusts other than those reported on Figures 3 and 4 may be precluded (see also Carreras and Druguet, 2014).Such extra thrusts have been drawn across the Axial Zone to absorb Alpine shortening, implying staking, internal deformation and northward (up to 90°) tilting of basement duplexes (Muñoz, 1992;Beaumont et al., 2000).The present study suggests instead that the Axial Zone is an asymmetrical Alpine pop-up, whose southern half has been flexed as a consequence of slip along curved, north-dipping thrust(s), forming an upward-fanning pattern of Variscan schistosity (e.g., Fig. 4, cross-section 3).The Alpine convergence record in the Pyrenees therefore needs to be substantially revised.

The Pyrenees within the closing Iberian-Armorican syntax
If the map development of the Cantabrian orocline is now well constrained and dated, its causes remain elusive (Gutiérrez-Alonso et al., 2012).Martínez-Catalán (2011) suggested that Variscan oroclines resulted from orogenparallel shortening within the late Paleozoic dextral shear corridor linking the Appalachians to the Urals defined by Arthaud and Matte (1977).Carreras and Druguet (2014) proposed that the Iberian-Armorican arc was a back fold developed in between large-scale en-échelon Riedel faults (especially the North Pyrenean fault) within that same dextral corridor.Whatever the far-field conditions, as a limb portion of the Cantabrian orocline, the Pyrenees developed a strain pattern that fits into the orocline closure scheme in a way that is summarized as follows.
Thin-skinned fold-and-thrust deformation attained the Pyrenees around 320-310 Ma, i.e., at a time HP nappe stacking (380-360 Ma) and exhumation (360-350 Ma) had long occurred in the hinterland of the orogen (Faure, 1995;Faure et al., 2008Faure et al., , 2009;;Matte, 2007;Martínez-Catalán et al., 2009;Ballèvre et al., 2009;2014;Fig.14a and b).The nappe stack became subject to partial melting and magmatism under a regime of dominant orogen-parallel flow that led to the emplacement of extensional gneiss domes and plutons (Fig. 14b).Later on (Fig. 14c), partial melting and lateral crustal flow came to affect a wider hinterland growing at the expense of its southern foreland that was recording active thrusting (e.g., Matte, 2007;Martínez-Catalán et al., 2009).Final amplification of the Iberian-Armorican syntax was recorded in its core by the closure of the Cantabrian orocline between 305 and 295 Ma (Weil et al., 2010).Coeval shortening and lateral flow of the Pyrenees from 310 to 300 Ma under an increasing component of transpression until at least 290 Ma responded to orocline closure.The Pyrenean strain field recorded lateral (i.e., limb-parallel) eastward escape of the crustal material pinched out in the inner syntax and extruded outward, whereas transpression recorded shortening across the orocline's limb (Fig. 14c).In this context, the dextral component of transpression in the Pyrenees has been imposed by the obliquity between overall N-S convergence and the NW-SE strike of the northern limb of the orocline (in Permian coordinates, Fig. 14c).The Pyrenees are peculiar among the HT terrains of the Variscan belt of SW Europe in the sense that the last deformation that left an imprint on the crustal architecture is compressional instead of being related to extensional collapse.,.The inset shows how syn-convergence lateral flow pattern of the Pyrenees fits into the frame of the closing syntax.Note that N-S extensional shearing in the North Pyrenean massifs may be younger (see text for further explanation).The Axial Zone of the Pyrenees has been kept attached to Iberia, which is restored to its Permian position.Main sources for map and kinematic data outside the Pyrenees: Faure et al. (2009), Ballèvre et al. (2014), Franke (2014), Lardeaux et al. (2014).Palinspastic position of Iberia is adapted from Carreras and Druguet (2014) and those of Corsica and Sardinia from Edel et al. (2014).
along the limb of the closing Cantabrian orocline.Orocline closure pinched off the crust enclosed in the inner syntax that escaped towards the SE (in Permian coordinates; Fig. 14c), providing a lower-strength, favorable boundary condition compared to the orocline's hinge.
N-S extensional exhumation of the granulitic crust in the North Pyrenean massifs does not readily fit into the kinematic framework described above (Fig. 14c).However, some of these massifs display longitudinal mineral stretching lineations (Fig. 3) and recent zircon U-Pb dating on the granulites suggests extension-driven cooling at high temperature significantly later than in the Axial Zone (300-280 Ma; Lemirre et al., 2016).Furthermore, the relative position of the North Pyrenean massifs and the Axial Zone on both sides of the North Pyrenean fault is not well constrained for pre-Cretaceous times.Therefore, although they possibly underwent early Axial Zone-like lateral flow, the North Pyrenean massifs preserve the record of N-S midto Late Permian extension comparable to that affecting pervasively the French Massif Central (Malavieille, 1993;Burg et al., 1994, Faure, 1995;Matte, 2007;Figs. 1 and 14c).Such a transverse extension may have been driven by collapse of a wide hinterland of higher topography, as opposed to transpression and lateral escape of the foreland Pyrenean crust from the closing orocline.Partitioning of stretching/extension directions between those two domains was allowed by the North Pyrenean fault.
The Pyrenean crust was an abnormally hot foreland undergoing regional HT/LP metamorphism between ca.310 and 300 Ma.Syn-convergence HT-LP metamorphism cannot be the sole consequence of the thermal maturation of an overthickened crust (Thompson and England, 1984), especially considering the absence of crustal-scale thrust and the lack of HP metamorphic relicts in the case of the Pyrenees.For that, an additional asthenospheric heat source is required in a context of attenuated or missing lithospheric mantle (Sandiford and Powell, 1991;Chardon et al., 2009).The switch from crustal to mantle and hybrid sources of 310-290 Ma plutonism in the Pyrenees (e.g., Kilzi et al., 2016) is in agreement with such a lithospheric configuration that enhances mantle-crust interactions.This switch, which is also documented across both limbs of the syntax after ca.330 Ma (Bussy et al., 2000;Gébelin et al., 2009;Gutiérrez-Alonso et al., 2011a, b;Couzinié et al., 2014), may be due to lithospheric mantle delamination (Rey et al., 1997;Gutiérrez-Alonso et al., 2011a, b;Laurent et al., 2017; see also the critical review of Martínez-Catalán et al., 2014).Franke (2014) argued that magmatism and HT-LP metamorphism were widespread over the entire orogen and its forelands before 330 Ma and suggested a "non orogenic" cause for a modification of the lithosphere such as mantle plumes.But pre-320 Ma zircon U-Pb and monazite U-Th-Pb ages from the vicinity of flysch basins are those of crustally-derived melts and have been challenged by recent geochronological reappraisals as exemplified in two key areas such as the Pyrenees (Denèle et al., 2014) and the Montagne Noire (Roger et al., 2015;Poujol et al., 2017;Fig. 1).
Whatever the cause(s) for the weakening of the lithospheric mantle under the Iberian -Armorican syntax (thermal erosion or delamination), a thin and hot lithosphere appears as a necessary condition for the 3D flow mode of the Pyrenean crust during orocline closure.A thin and hot lithosphere is also required for orocline bending because a stiff foreland lithosphere that may not have been bent otherwise.

Conclusion
The specificity of the Pyrenean Variscan crust is to have evolved from a cold foreland setting to a hot orogenic setting without having undergone typical mountain building processes in the meantime (crustal-scale thrust nappe tectonics, HP metamorphism).As a thin and hot portion of continental lithosphere, the Pyrenean domain has been involved in the closure of an orogenic syntax by developing strain and metamorphic patterns typical of hot orogens.These patterns reflect the competition between upper crustal thickening and lateral flow of a weak and buoyant lower crust within the kinematic framework of a closing syntax.
An attachment has acted as a structural buffer between laterally flowing lower crust and transpressional thickening of the upper crust of the Variscan Pyrenees.This attachment corresponds to a temperature-controlled rheological threshold, which also allowed for the formation of transtensional gneiss domes at the roof of the lower crust during convergence.Transtensional unroofing of the domes into the upper crust involved a combination of forced lateral flow and buoyancy of the partially molten lower crust.Late shortening-induced buckling of the domes during strain localization in steep crustal-scale transpressive shear zones attests to an increasing coupling between the lower crust and the upper crust during regional cooling.The combination of dextral transpression and eastward flow in the Pyrenees results from the shortening and lateral escape of hot and weak crust along the inner northern limb of the closing Cantabrian orocline between ca.305 and 295 Ma.The conditions for the switch from crust-to mantlederived magmatism in the Iberian-Armorican arc, orocline closure and escape of hot foreland crust from the syntax are fulfilled in a context of thin and hot lithosphere having been delaminated or thermally eroded.

Fig. 1 .
Fig. 1.Sketch structural map of the Variscan belt of Europe showing the main outcrops of Pre-Mesozoic basement (in grey).Iberia and the Pyrenees as well as Sardinia and Corsica are shown in their current position.AM: Armorican Massif; BM: Bohemian Massif; FMC: French Massif Central; IM: Iberian Massif. Fig.8

Fig. 2 .
Fig. 2. Sketch map of the Paleozoic crust of the Pyrenees showing the location and name of the geological features discussed in the text.Domes and plutons are listed from West to East.Locations of the cross-sections of Figure 4 (labeled 1 to 5) and that of Figure 12a are shown.

Fig. 3 .
Fig. 3. (a) Structural and kinematic map of the Paleozoic crust of the Pyrenees.The Basque Massifs are displaced from their original position (see Fig. 2).The reader may refer to Figure 2 for the nomenclature of the main structural features.(b) Synthetic structural map.Transverse Li lineation trajectories have been drawn only where those lineations are dominant over large portions of the infrastructure.SeeDebon and Guitard (1996) for the corresponding regional-scale isograd map pattern.

Foliation
ig. 5. Planar and linear fabric measurements from the Axial Zone.(a) Ss schistosity poles of the superstructure.(b) Ls mineral-stretching lineations of the superstructure.Ss/Ls fabrics were statistically steeper than they appear before post-Paleozoic flexure of the southern half of the Axial Zone (e.g., Fig.4).(c) Si foliation poles of the infrastructure.(d) Li mineral-stretching lineations of the infrastructure.The red dot represents the mean pole of Ss schistosity and the red star is the pole to the planar best fit of the Si foliations, which is interpreted as the domes' hinge trend.Dark blue dots represent mean mineral-stretching lineations.Measurements are too numerous to be shown in (a) and (c).Contours are 2s.The data include those acquired for the present study and those collected from maps and the literature (see Supplementary Material).Lower hemisphere, equal area projection.

Fig. 9 .
Fig. 9. Block diagram of the southeastern Bossost dome and its envelope (see Fig. 8 for location and color codes).See text for further explanation.
Fig. 11.(a) Schematic 3D model of the Lys-Caillaouas massif.Compiled from maps and cross-sections ofClin et al. (1986Clin et al. ( , 1989)),Kriegsman et al. (1989), Hilario-Or us (2004)  and the present work (see Figure3afor kinematic information in map view).Alpine cleavage along the Esera-Gistain fault, which reactivates the shear zone, is omitted.(b) Normal-dextral C/S structures observed in the diorite along the southern boundary shear zone, indicating top-to-the ESE shear sense (the foliation dips ca.40°to the South).(c) Plane view of C' shear bands in sillimanite micaschists against the deformed plutonic rocks (sinistral sense of shear).Thin-sections are cut in XZ planes of the deformation ellipsoid.Mineral abbreviations fromKretz (1983).

Fig. 12 .
Fig. 12.(a) Cross-section at the Western termination of the Axial Zone, preserving early Variscan (D1) structures.See Figure 2 for location of the cross-section.(b) Southwestward verging folds with axial planar cleavage and top-to-the W reverse shears, Namurian flysch (outcrop located a few kilometers north of the cross-section).
Block-diagram summarizing two stages in the late-Variscan crustal evolution of the Axial Zone of the Pyrenees.See text for further explanations.

AFig. 14 .
Fig. 14.Three-stage reconstruction of the Iberian-Armorican arc during the Carboniferous and the Permian.(a) Late collision stage at ca. 330 Ma.(b) Syn-convergence extension of the hinterland and fold-and-thrust tectonics in the southern foreland.(c) End of orocline bending, intra-syntax HT deformation (ca.310-290 Ma).The inset shows how syn-convergence lateral flow pattern of the Pyrenees fits into the frame of the closing syntax.Note that N-S extensional shearing in the North Pyrenean massifs may be younger (see text for further explanation).The Axial Zone of the Pyrenees has been kept attached to Iberia, which is restored to its Permian position.Main sources for map and kinematic data outside the Pyrenees:Faure et al. (2009),Ballèvre et al. (2014),Franke (2014),Lardeaux et al. (2014).Palinspastic position of Iberia is adapted fromCarreras and Druguet (2014) and those of Corsica and Sardinia fromEdel et al. (2014).